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The Defender has a standard remote vehicle starting system, so the vehicle can be started from inside the driver's house. This allows the driver to comfortably warm up the engine before going out to the vehicle. The climate system will also automatically heat or cool the interior. The 4Runner doesn’t offer a remote starting system.
Unlike the driver-only memory system in the 4Runner Premium/Limited/Trailhunter/TRD Pro/Platinum, the Defender has standard driver and passenger memory, so that when drivers switch, the memory setting adjusts the driver’s seat, steering wheel position, outside mirror angle and climate settings and the front passenger seat also adjusts to the new passenger’s preset preferences.
The Defender’s standard Keyless Entry allows you to unlock the doors from any outside door handle, unlock the cargo door, and start the engine, all without removing the key from the pocket or purse. Smart Key standard on the 4Runner doesn’t offer a sensor on the rear doors, so you’ll have to reach a front handle to unlock the rear door.
The Defender’s rain-sensitive wipers adjust their speed and turn on and off automatically based on the amount of rainfall on the windshield. This allows the driver to concentrate on driving without constantly adjusting the wipers. The 4Runner’s standard manually variable intermittent wipers have to be constantly adjusted.
Heated windshield washer nozzles are optional on the Defender to prevent washer fluid and nozzles from freezing and help continue to keep the windshield clear in sub-freezing temperatures. The 4Runner doesn’t offer heated windshield washer nozzles.
In poor weather, headlights can lose their effectiveness as grime builds up on their lenses. This can reduce visibility without the driver realizing. The Defender offers available headlight washers to keep headlight output high. The 4Runner doesn’t offer headlight washers.
The Defender’s standard rear and side view mirrors have an automatic dimming feature. These mirrors can be set to automatically darken quickly when headlights shine on them, keeping following vehicles from blinding or distracting the driver. The 4Runner offers an automatic rear view mirror, but its side mirrors don’t dim.
The Land Rover Defender has power folding mirrors that offer unparalleled convenience while maneuvering in tight places or parked. With the mirrors folded it’s easier to walk past and help prevent parking damage. The mirrors can be quickly unfolded when the driver is ready to leave. Power folding mirrors are only offered on the 4Runner Limited/Trailhunter/TRD Pro/Platinum.
The Defender has standard heated front and optional heated second and third row seats, which keep the driver and passengers extremely comfortable in the winter. Toyota doesn’t offer heated seats in the third row of the 4Runner.
Optional air conditioned front and second row seats keep the Defender’s passengers comfortable and take the sting out of hot leather in summer. The 4Runner doesn’t offer air-conditioned seats for the second row.
The Defender has a standard heated steering wheel to take the chill out of steering on extremely cold winter days before the vehicle heater warms up. A heated steering wheel is only available on the 4Runner Premium/Limited/Platinum/Trailhunter/TRD Pro.
The Defender has a standard dual zone air conditioning allows the driver and front passenger to choose two completely different temperatures so people with different temperature preferences won’t have to compromise. This makes both the driver and front passenger as comfortable as possible. Dual zone air conditioning is only available on the 4Runner Limited/Platinum/Trailhunter/TRD Pro.
The Land Rover Defender has a standard Homelink wireless remote control system for garage door operation and device management, conveniently located on the rear view mirror. Homelink® eliminates the need for separate garage door openers and associated risks of losing, breaking, or having dead batteries. Homelink® is only available on the 4Runner Premium/Limited/Platinum/Trailhunter/TRD Pro.
To quickly and conveniently keep personal devices charged without cables tangling and wearing out, the Land Rover Defender has a standard wireless phone charging system (Qi) in the center console. Only the 4Runner TRD/Limited/Platinum/Trailhunter offers wireless charging.
