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Compare the2025 Land Rover DefenderVS 2025 Toyota 4Runner

2025 Land Rover Defender
2025 Toyota 4Runner

Safety

© 1999 - 2025Advanta-STAR Automotive Research, all rights reserved. This vehicle comparison and all of the content in it are provided only by license from Advanta-STAR Automotive Research Corporation of America (“Advanta-STAR”). If you are not a legally licensed user of this vehicle comparison, it is against federal law to access it, copy it, forward it, or use it in any manner whatsoever. Any unauthorized use of this vehicle comparison is a violation of U.S. and international law and is punishable criminally and civilly. Removal of this watermark/notification without prior written license and approval received from Advanta-STAR is an agreement, understanding, and/or stipulation by the person(s), entities, agents, attorneys, and any other persons involved in the removal of this watermark/notification (including but not limited to Search Optics, LLC and any and all parent entities, sister entities, and subsidiary entities of Search Optics, LLC and/or any other entity, agent, attorney, and persons related in any manner to Search Optics, LLC) to: 1) an agreed upon amount of liquidated monetary damages of a minimum of $1,250,000.00 US Dollars in favor of Advanta-STAR; 2) the jurisdiction and enforcement of any legal claims associated with this matter asserted by Advanta-STAR in the United States Federal District Court in Portand, Oregon; and 3) service of process of any legal claims asserted by Advanta-STAR associated with this matter may be accomplished by First-Class Postage by the United States Postal Service or comparable service. XPYNN-M34HG 2a06:98c0:3600::103 2025/05/30

For enhanced safety, the front and second-row seat shoulder belts of the Land Rover Defender have pretensioners to tighten the seatbelts and eliminate dangerous slack in the event of a collision and force limiters to limit the pressure the belts will exert on the passengers. The Toyota 4Runner doesn’t offer pretensioners for its second-row seat belts.

Both the Defender 110/130 and 4Runner have child safety locks to prevent children from opening the rear doors. The Defender has power child safety locks, allowing the driver to activate and deactivate them from the driver's seat and to know when they're engaged. The 4Runner’s child locks have to be individually engaged at each rear door with a manual switch. The driver can’t know the status of the locks without opening the doors and checking them.

To provide maximum traction and stability on all roads, All-Wheel Drive is standard on the Defender. But it costs extra on the 4Runner.

The Land Rover Defender’s rear backup camera has a standard washer for maintaining a clear view under various conditions. In contrast, the Toyota 4Runner does not offer a rear camera washer, meaning its effectiveness relies on manual cleaning by the user when necessary.

To help make backing out of a parking space safer, the Defender has standard Rear Traffic Monitor with Rear Traffic Braking, systems which detect vehicles approaching from the sides and can automatically apply the brakes to prevent a collision. Only the 4Runner Limited/Platinum/Trailhunter/TRD Pro offers Parking Support Brake.

For better protection of the passenger compartment, the Defender uses safety cell construction with a three-dimensional high-strength frame that surrounds the passenger compartment. It provides extra impact protection and a sturdy mounting location for door hardware and side impact beams. The 4Runner uses a body-on-frame design, which has no frame members above the floor of the vehicle.

Both the Defender and the 4Runner have standard driver and passenger frontal airbags, front side-impact airbags, side-impact head airbags, four-wheel antilock brakes, traction control, electronic stability systems to prevent skidding, crash mitigating brakes, daytime running lights, lane departure warning systems, blind spot warning systems, rearview cameras, rear cross-path warning and driver alert monitors.

Warranty

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The Defender comes with a full 4-year/50,000-mile basic warranty, which covers the entire truck and includes 24-hour roadside assistance. The 4Runner’s 3-year/36,000-mile basic warranty expires 1 year or 14,000 miles sooner.

The Defender’s corrosion warranty is 1 year longer than the 4Runner’s (6 vs. 5 years).

Engine

© 1999 - 2025Advanta-STAR Automotive Research, all rights reserved. This vehicle comparison and all of the content in it are provided only by license from Advanta-STAR Automotive Research Corporation of America (“Advanta-STAR”). If you are not a legally licensed user of this vehicle comparison, it is against federal law to access it, copy it, forward it, or use it in any manner whatsoever. Any unauthorized use of this vehicle comparison is a violation of U.S. and international law and is punishable criminally and civilly. Removal of this watermark/notification without prior written license and approval received from Advanta-STAR is an agreement, understanding, and/or stipulation by the person(s), entities, agents, attorneys, and any other persons involved in the removal of this watermark/notification (including but not limited to Search Optics, LLC and any and all parent entities, sister entities, and subsidiary entities of Search Optics, LLC and/or any other entity, agent, attorney, and persons related in any manner to Search Optics, LLC) to: 1) an agreed upon amount of liquidated monetary damages of a minimum of $1,250,000.00 US Dollars in favor of Advanta-STAR; 2) the jurisdiction and enforcement of any legal claims associated with this matter asserted by Advanta-STAR in the United States Federal District Court in Portand, Oregon; and 3) service of process of any legal claims asserted by Advanta-STAR associated with this matter may be accomplished by First-Class Postage by the United States Postal Service or comparable service. XPYNN-M34HG 2a06:98c0:3600::103 2025/05/30

The Defender has more powerful engines than the 4Runner:

Horsepower

Torque

Defender P300 2.0 turbo 4-cylinder

296 HP

295 lbs.-ft.

Defender P400 3.0 turbo/supercharged 6-cylinder hybrid

395 HP

406 lbs.-ft.

Defender P500 5.0 supercharged V8

493 HP

450 lbs.-ft.

Defender P525 5.0 supercharged V8

518 HP

461 lbs.-ft.

Defender OCTA 4.4 turbo V8 hybrid

626 HP

553 lbs.-ft.

4Runner 2.4 turbo 4-cylinder

278 HP

317 lbs.-ft.

4Runner Trailhunter 2.4 turbo 4-cylinder hybrid

323 HP

465 lbs.-ft.

4Runner 2.4 turbo 4-cylinder hybrid

326 HP

465 lbs.-ft.

As tested in Car and Driver the Defender P400 3.0 turbo/supercharged 6-cylinder hybrid is faster than the 4Runner Trailhunter 2.4 turbo 4-cylinder hybrid:

Defender

4Runner

Zero to 60 MPH

6.2 sec

6.7 sec

5 to 60 MPH Rolling Start

6.8 sec

7.4 sec

Passing 30 to 50 MPH

4 sec

4.1 sec

Passing 50 to 70 MPH

4.5 sec

4.8 sec

Quarter Mile

14.6 sec

15.3 sec

Speed in 1/4 Mile

95 MPH

88 MPH

Fuel Economy and Range

© 1999 - 2025Advanta-STAR Automotive Research, all rights reserved. This vehicle comparison and all of the content in it are provided only by license from Advanta-STAR Automotive Research Corporation of America (“Advanta-STAR”). If you are not a legally licensed user of this vehicle comparison, it is against federal law to access it, copy it, forward it, or use it in any manner whatsoever. Any unauthorized use of this vehicle comparison is a violation of U.S. and international law and is punishable criminally and civilly. Removal of this watermark/notification without prior written license and approval received from Advanta-STAR is an agreement, understanding, and/or stipulation by the person(s), entities, agents, attorneys, and any other persons involved in the removal of this watermark/notification (including but not limited to Search Optics, LLC and any and all parent entities, sister entities, and subsidiary entities of Search Optics, LLC and/or any other entity, agent, attorney, and persons related in any manner to Search Optics, LLC) to: 1) an agreed upon amount of liquidated monetary damages of a minimum of $1,250,000.00 US Dollars in favor of Advanta-STAR; 2) the jurisdiction and enforcement of any legal claims associated with this matter asserted by Advanta-STAR in the United States Federal District Court in Portand, Oregon; and 3) service of process of any legal claims asserted by Advanta-STAR associated with this matter may be accomplished by First-Class Postage by the United States Postal Service or comparable service. XPYNN-M34HG 2a06:98c0:3600::103 2025/05/30

Regardless of its engine, the Defender’s engine automatically turns off when the vehicle is stopped, saving fuel and reducing pollution. If the conditions warrant or the driver wishes, the system can be manually disabled at any time for the duration of a trip. Toyota only offers an automatic engine start/stop system on the 4Runner Limited/i-FORCE MAX.

The Defender has 4.8 gallons more fuel capacity than the 4Runner (23.8 vs. 19 gallons), for longer range between fill-ups.

Brakes and Stopping

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For better stopping power the Defender’s brake rotors are larger than those on the 4Runner:

Defender

Defender V8

4Runner

Front Rotors

14.3 inches

14.9 inches

13.4 inches

Rear Rotors

13.8 inches

14.3 inches

13.2 inches

The Defender stops much shorter than the 4Runner:

Defender

4Runner

70 to 0 MPH

167 feet

178 feet

Car and Driver

60 to 0 MPH

120 feet

127 feet

Motor Trend

Tires and Wheels

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For better traction, the Defender’s optional tires are larger than the largest tires available on the 4Runner (275/45R22 vs. 265/70R17).

The Defender’s standard tires provide better handling because they have a lower 65 series profile (height to width ratio) that provides a stiffer sidewall than the 4Runner SR5’s standard 70 series tires. The Defender’s optional tires have a lower 45 series profile than the 4Runner TRD Sport/Limited/Platinum’s 55 series tires.

For better ride, handling and brake cooling the Defender has standard 19-inch wheels. Smaller 17-inch wheels are standard on the 4Runner SR5. The Defender’s optional 22-inch wheels are larger than the 20-inch wheels on the 4Runner TRD Sport/Limited/Platinum.

Suspension and Handling

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For a smoother ride and more stable handling, the Defender 110’s wheelbase is 6.8 inches longer than on the 4Runner (119 inches vs. 112.2 inches).

For better handling and stability, the average track (width between the wheels) on the Defender is 1.3 inches wider in the front and 1.1 inches wider in the rear than the average track on the 4Runner.

The Defender 130 Outbound handles at .77 G’s, while the 4Runner Trailhunter pulls only .74 G’s of cornering force in a Car and Driver skidpad test.

The Defender 110 V8 executes Motor Trend’s “Figure Eight” maneuver 1 seconds quicker than the 4Runner TRD Off-Road Premium (27 seconds @ .66 average G’s vs. 28 seconds @ .6 average G’s).

For better maneuverability, the Defender 90’s turning circle is 1.4 feet tighter than the 4Runner’s (38 feet vs. 39.4 feet).

For greater off-road capability the Defender 90 has a greater minimum ground clearance than the 4Runner SR5 (8.9 vs. 8.1 inches), allowing the Defender to travel over rougher terrain without being stopped or damaged. The Defender 110 OCTA’s minimum ground clearance is 2.6 inches higher than on the 4Runner Trailhunter/TRD Pro (12.7 vs. 10.1 inches).

Chassis

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Unibody construction lowers the Defender’s center of gravity significantly without reducing ground clearance. This contributes to better on the road handling and better off-road performance and stability. In addition, unibody construction makes the chassis stiffer, improving handling and reducing squeaks and rattles. The 4Runner doesn’t use unibody construction, but a body-on-frame design.

As tested by Car and Driver while cruising at 70 MPH, the interior of the Defender 130 Outbound is quieter than the 4Runner Trailhunter (67 vs. 70 dB).

Passenger Space

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The Defender 130 offers optional seating for 8 passengers; the 4Runner can only carry up to 7.

Cargo Capacity

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The Defender 130’s cargo area provides more volume than the 4Runner.

Defender

4Runner

Behind Third Seat

15.3 cubic feet

12.1 cubic feet

Second Seat Folded

89 cubic feet

84.4 cubic feet

Towing

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The Defender’s standard towing capacity is much higher than the 4Runner’s (7716 vs. 6000 pounds). Maximum trailer towing in the Toyota 4Runner is only 6000 pounds. The Defender offers up to a 8201 lbs. towing capacity.

Ergonomics

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The Defender has a standard remote vehicle starting system, so the vehicle can be started from inside the driver's house. This allows the driver to comfortably warm up the engine before going out to the vehicle. The climate system will also automatically heat or cool the interior. The 4Runner doesn’t offer a remote starting system.

Unlike the driver-only memory seat and mirrors in the 4Runner Premium/Limited/Trailhunter/TRD Pro/Platinum, the Defender offers an optional driver and passenger memory, so that when drivers switch, the memory setting adjusts the driver’s seat, steering wheel position, outside mirror angle and climate settings and the front passenger seat also adjusts to the new passenger’s preset preferences.

The Defender’s optional easy entry system raises the steering wheel and glides the driver’s seat back when the door is unlocked or the ignition is switched off, making it easier for the driver to get in and out. The 4Runner doesn’t offer an easy entry system.

The Defender’s standard Keyless Entry allows you to unlock the doors from any outside door handle, unlock the cargo door, and start the engine, all without removing the key from the pocket or purse. Smart Key standard on the 4Runner doesn’t offer a sensor on the cargo door or the rear doors, so you’ll have to reach a front handle to unlock the rear door.

The Defender’s rain-sensitive wipers adjust their speed and turn on and off automatically based on the amount of rainfall on the windshield. This allows the driver to concentrate on driving without constantly adjusting the wipers. The 4Runner’s standard manually variable intermittent wipers have to be constantly adjusted.

Heated windshield washer nozzles are optional on the Defender to prevent washer fluid and nozzles from freezing and help continue to keep the windshield clear in sub-freezing temperatures. The 4Runner doesn’t offer heated windshield washer nozzles.

To improve rear visibility by keeping the rear window clear, the Defender has a standard rear fixed intermittent wiper with a full on position. The rear wiper standard on the 4Runner only has an intermittent setting, so in a hard rain visibility isn’t as good.

The Defender’s standard rear and side view mirrors have an automatic dimming feature. These mirrors can be set to automatically darken quickly when headlights shine on them, keeping following vehicles from blinding or distracting the driver. The 4Runner offers an automatic rear view mirror, but its side mirrors don’t dim.

The Land Rover Defender has power folding mirrors that offer unparalleled convenience while maneuvering in tight places or parked. With the mirrors folded it’s easier to walk past and help prevent parking damage. The mirrors can be quickly unfolded when the driver is ready to leave. Power folding mirrors are only offered on the 4Runner Limited/Trailhunter/TRD Pro/Platinum.

The Defender has a standard heated steering wheel to take the chill out of steering on extremely cold winter days before the vehicle heater warms up. A heated steering wheel is only available on the 4Runner Premium/Limited/Platinum/Trailhunter/TRD Pro.

The Defender has a standard dual zone air conditioning allows the driver and front passenger to choose two completely different temperatures so people with different temperature preferences won’t have to compromise. This makes both the driver and front passenger as comfortable as possible. Dual zone air conditioning is only available on the 4Runner Limited/Platinum/Trailhunter/TRD Pro.

The Land Rover Defender has a standard Homelink wireless remote control system for garage door operation and device management, conveniently located on the rear view mirror. Homelink® eliminates the need for separate garage door openers and associated risks of losing, breaking, or having dead batteries. Homelink® is only available on the 4Runner Premium/Limited/Platinum/Trailhunter/TRD Pro.

To quickly and conveniently keep personal devices charged without cables tangling and wearing out, the Land Rover Defender has a standard wireless phone charging system (Qi) in the center console. Only the 4Runner TRD/Limited/Platinum/Trailhunter offers wireless charging.

Model Availability

© 1999 - 2025Advanta-STAR Automotive Research, all rights reserved. This vehicle comparison and all of the content in it are provided only by license from Advanta-STAR Automotive Research Corporation of America (“Advanta-STAR”). If you are not a legally licensed user of this vehicle comparison, it is against federal law to access it, copy it, forward it, or use it in any manner whatsoever. Any unauthorized use of this vehicle comparison is a violation of U.S. and international law and is punishable criminally and civilly. Removal of this watermark/notification without prior written license and approval received from Advanta-STAR is an agreement, understanding, and/or stipulation by the person(s), entities, agents, attorneys, and any other persons involved in the removal of this watermark/notification (including but not limited to Search Optics, LLC and any and all parent entities, sister entities, and subsidiary entities of Search Optics, LLC and/or any other entity, agent, attorney, and persons related in any manner to Search Optics, LLC) to: 1) an agreed upon amount of liquidated monetary damages of a minimum of $1,250,000.00 US Dollars in favor of Advanta-STAR; 2) the jurisdiction and enforcement of any legal claims associated with this matter asserted by Advanta-STAR in the United States Federal District Court in Portand, Oregon; and 3) service of process of any legal claims asserted by Advanta-STAR associated with this matter may be accomplished by First-Class Postage by the United States Postal Service or comparable service. XPYNN-M34HG 2a06:98c0:3600::103 2025/05/30

The Land Rover Defender comes in two door and four door bodystyles; the Toyota 4Runner isn’t available as a two door.

Recommendations

© 1999 - 2025Advanta-STAR Automotive Research, all rights reserved. This vehicle comparison and all of the content in it are provided only by license from Advanta-STAR Automotive Research Corporation of America (“Advanta-STAR”). If you are not a legally licensed user of this vehicle comparison, it is against federal law to access it, copy it, forward it, or use it in any manner whatsoever. Any unauthorized use of this vehicle comparison is a violation of U.S. and international law and is punishable criminally and civilly. Removal of this watermark/notification without prior written license and approval received from Advanta-STAR is an agreement, understanding, and/or stipulation by the person(s), entities, agents, attorneys, and any other persons involved in the removal of this watermark/notification (including but not limited to Search Optics, LLC and any and all parent entities, sister entities, and subsidiary entities of Search Optics, LLC and/or any other entity, agent, attorney, and persons related in any manner to Search Optics, LLC) to: 1) an agreed upon amount of liquidated monetary damages of a minimum of $1,250,000.00 US Dollars in favor of Advanta-STAR; 2) the jurisdiction and enforcement of any legal claims associated with this matter asserted by Advanta-STAR in the United States Federal District Court in Portand, Oregon; and 3) service of process of any legal claims asserted by Advanta-STAR associated with this matter may be accomplished by First-Class Postage by the United States Postal Service or comparable service. XPYNN-M34HG 2a06:98c0:3600::103 2025/05/30

Motor Trend selected the Defender as their 2021 Sport Utility of the Year. The 4Runner has never been chosen.

A group of representative automotive journalists from North America selected the Defender as the 2021 North American Utility Vehicle of the Year. The 4Runner has never been chosen.

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