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The engine computer on the Wrangler automatically engages the starter until the car starts with one twist of the key and disables the starter while the engine is running. The 4Runner’s starter can be accidentally engaged while the engine is running, making a grinding noise and possibly damaging the starter and ring gear.
The Wrangler’s instruments include an oil pressure gauge and a temperature gauge - which could save your engine! Often ‘idiot lights’ don’t warn you until damage has been done. The 4Runner does not have an oil pressure gauge.
The power windows available on both the Wrangler and the 4Runner have locks to prevent small children from operating them. When the lock on the Wrangler is engaged the driver can still operate all of the windows, for instance to close one opened by a child. The 4Runner prevents the driver from operating the other windows just as it does the other passengers.
The Wrangler’s power window, power lock, power mirror and cruise control switches are lit from behind, making them plainly visible and easier to operate at night. The 4Runner’s power mirror and cruise control switches are unlit, making them difficult to find at night and operate safely.
The Wrangler’s speed-sensitive wipers speed up when the vehicle does, so that the driver doesn’t have to continually adjust the speed of the wipers. The 4Runner’s manually variable intermittent wipers have to be constantly adjusted.
The Wrangler’s optional power mirror controls are mounted on the door for easy access. The 4Runner’s power mirror controls are on the dash where they are possibly hidden by the steering wheel and are awkward to manipulate.