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The Discovery has a standard remote vehicle starting system, so the vehicle can be started from inside the driver's house. This allows the driver to comfortably warm up the engine before going out to the vehicle. The climate system will also automatically heat or cool the interior. The 4Runner doesn’t offer a remote starting system.
The engine computer on the Discovery disables the starter while the engine is running. The 4Runner’s starter can be accidentally engaged while the engine is running, making a grinding noise and possibly damaging the starter and ring gear.
Unlike the driver-only memory seat in the 4Runner Limited, the Discovery offers an optional driver and passenger memory, so that when drivers switch, the memory setting adjusts the driver’s seat, steering wheel position and outside mirror angle and the front passenger seat also adjusts to the new passenger’s preset preferences.
The Discovery’s optional easy entry system glides the driver’s seat back when the door is unlocked or the ignition is switched off, making it easier for the driver to get in and out. The 4Runner doesn’t offer an easy entry system.
The Discovery offers an optional heads-up display that projects speed and navigation instruction readouts in front of the driver’s line of sight, allowing drivers to view information without diverting their eyes from the road. The 4Runner doesn’t offer a heads-up display.
The Discovery’s power parking brake sets with one touch and releases with one touch or automatically. The 4Runner’s parking brake has to be released manually.
The power windows standard on both the Discovery and the 4Runner have locks to prevent small children from operating them. When the lock on the Discovery is engaged the driver can still operate all of the windows, for instance to close one opened by a child. The 4Runner prevents the driver from operating the other windows just as it does the other passengers.
The Discovery has a standard locking fuel door with a remote release located convenient to the driver. A locking fuel door helps prevent fuel theft and vandalism, such as sugar in the tank. The 4Runner doesn’t offer a locking fuel door.
The Discovery’s rain-sensitive wipers adjust their speed and turn on and off automatically based on the amount of rainfall on the windshield. This allows the driver to concentrate on driving without constantly adjusting the wipers. The 4Runner’s manually variable intermittent wipers have to be constantly adjusted.
Heated windshield washer nozzles are optional on the Discovery to prevent washer fluid and nozzles from freezing and help continue to keep the windshield clear in sub-freezing temperatures. The 4Runner doesn’t offer heated windshield washer nozzles.
To improve rear visibility by keeping the rear window clear, the Discovery has a standard rear fixed intermittent wiper with a full on position. The rear wiper standard on the 4Runner only has an intermittent setting, so in a hard rain visibility isn’t as good.
In poor weather, headlights can lose their effectiveness as grime builds up on their lenses. This can reduce visibility without the driver realizing. The Discovery HSE/Metropolitan offers available headlight washers to keep headlight output high. The 4Runner doesn’t offer headlight washers.
The Discovery has a standard automatic headlight on/off feature. When the ignition is on, the headlights automatically turn on at dusk and off after dawn. When the ignition turns off, the headlights turn off after a delay timed to allow you to securely get to your front door. The 4Runner has an automatic headlight on/off feature standard only on the Limited/TRD Pro.
The Discovery’s power mirror controls are mounted on the door for easy access. The 4Runner’s power mirror controls are on the dash where they are possibly hidden by the steering wheel and are awkward to manipulate.
When the Discovery with available tilt-down mirrors is put in reverse, both rearview mirrors tilt from their original position. This gives the driver a better view of the curb during parallel parking maneuvers. Shifting out of reverse puts the mirrors into their original positions. The 4Runner’s mirrors don’t automatically adjust for backing.
The Discovery’s standard rear and side view mirrors have an automatic dimming feature. These mirrors can be set to automatically darken quickly when headlights shine on them, keeping following vehicles from blinding or distracting the driver. The 4Runner offers an automatic rear view mirror, but its side mirrors don’t dim.
The Discovery has standard heated front seats. Heated front seats are only available on the 4Runner Premium/Limited/TRD Pro. The Discovery also offers optional heated second and third row seats to keep those passengers extremely comfortable in the winter. Neither heated second nor third row seats are available in the 4Runner.
The Discovery HSE/Metropolitan has standard front air conditioned seats and the Discovery offers them optionally in the second row. This keeps the passengers comfortable and takes the sting out of hot seats in summer. The 4Runner doesn’t offer air-conditioned seats for the second row.
On extremely cold winter days, the Discovery’s optional heated steering wheel provides comfort, allowing the driver to steer safely and comfortably before the vehicle heater warms up. The 4Runner doesn’t offer a heated steering wheel.
The Discovery HSE/Metropolitan offers optional massaging front seats in order to maximize comfort and eliminate fatigue on long trips. Massaging seats aren’t available in the 4Runner.
The Discovery has a standard dual zone air conditioning allows the driver and front passenger to choose two completely different temperatures so people with different temperature preferences won’t have to compromise. This makes both the driver and front passenger as comfortable as possible. Dual zone air conditioning is only available on the 4Runner Limited/TRD Pro.
To quickly and conveniently keep personal devices charged without cables tangling and wearing out, the Land Rover Discovery has a standard wireless phone charging system (Qi) in the center console. The 4Runner doesn’t offer wireless personal charging.
The Discovery’s Park Assist can parallel park or back into a parking spot by itself, with the driver only controlling speed with the brake pedal. The 4Runner doesn’t offer an automated parking system.