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The Defender has a standard remote vehicle starting system, so the vehicle can be started from inside the driver's house. This allows the driver to comfortably warm up the engine before going out to the vehicle. The climate system will also automatically heat or cool the interior. The 4Runner doesn’t offer a remote starting system.
The engine computer on the Defender automatically engages the starter until the car starts with one twist of the key and disables the starter while the engine is running. The 4Runner’s starter can be accidentally engaged while the engine is running, making a grinding noise and possibly damaging the starter and ring gear.
Unlike the driver-only memory seat in the 4Runner Limited, the Defender (except 110 Base) has standard driver and passenger memory, so that when drivers switch, the memory setting adjusts the driver’s seat, steering wheel position (with optional power wheel adjuster), outside mirror angle and climate settings and the front passenger seat also adjusts to the new passenger’s preset preferences.
The Defender (except 110 Base)’s standard easy entry system raises the steering wheel and glides the driver’s seat back when the door is unlocked or the ignition is switched off, making it easier for the driver to get in and out. The 4Runner doesn’t offer an easy entry system.
The Defender (except Base) offers an available heads-up display that projects speed and navigation instruction readouts in front of the driver’s line of sight, allowing drivers to view information without diverting their eyes from the road. The 4Runner doesn’t offer a heads-up display.
The Defender’s power parking brake sets with one touch and releases with one touch or automatically. The 4Runner’s parking brake has to be released manually.
The power windows standard on both the Defender and the 4Runner have locks to prevent small children from operating them. When the lock on the Defender is engaged the driver can still operate all of the windows, for instance to close one opened by a child. The 4Runner prevents the driver from operating the other windows just as it does the other passengers.
The Defender has a standard locking fuel door with a remote release located convenient to the driver. A locking fuel door helps prevent fuel theft and vandalism, such as sugar in the tank. The 4Runner doesn’t offer a locking fuel door.
The Defender’s rain-sensitive wipers adjust their speed and turn on and off automatically based on the amount of rainfall on the windshield. This allows the driver to concentrate on driving without constantly adjusting the wipers. The 4Runner’s manually variable intermittent wipers have to be constantly adjusted.
Heated windshield washer nozzles are optional on the Defender to prevent washer fluid and nozzles from freezing and help continue to keep the windshield clear in sub-freezing temperatures. The 4Runner doesn’t offer heated windshield washer nozzles.
To improve rear visibility by keeping the rear window clear, the Defender has a standard rear fixed intermittent wiper with a full on position. The rear wiper standard on the 4Runner only has an intermittent setting, so in a hard rain visibility isn’t as good.
In poor weather, headlights can lose their effectiveness as grime builds up on their lenses. This can reduce visibility without the driver realizing. The Defender offers available headlight washers to keep headlight output high. The 4Runner doesn’t offer headlight washers.
The Defender has a standard automatic headlight on/off feature. When the ignition is on, the headlights automatically turn on at dusk and off after dawn. When the ignition turns off, the headlights turn off after a delay timed to allow you to securely get to your front door. The 4Runner has an automatic headlight on/off feature standard only on the Limited/TRD Pro.
The Defender’s power mirror controls are mounted on the armrest for easy access. The 4Runner’s power mirror controls are on the dash where they are possibly hidden by the steering wheel and are awkward to manipulate.
When the Defender is put in reverse, both rearview mirrors tilt from their original position. This gives the driver a better view of the curb during parallel parking maneuvers. Shifting out of reverse puts the mirrors into their original positions. The 4Runner’s mirrors don’t automatically adjust for backing.
The Defender’s standard rear and side view mirrors have an automatic dimming feature. These mirrors can be set to automatically darken quickly when headlights shine on them, keeping following vehicles from blinding or distracting the driver. The 4Runner offers an automatic rear view mirror, but its side mirrors don’t dim.
The Defender has standard heated front seats. Heated front seats are only available on the 4Runner Premium/Limited/TRD Pro. The Defender also offers optional heated second and third row seats to keep those passengers extremely comfortable in the winter. Neither heated second nor third row seats are available in the 4Runner.
Optional air conditioned front and second row seats keep the Defender’s passengers comfortable and take the sting out of hot leather in summer. The 4Runner doesn’t offer air-conditioned seats for the second row.
On extremely cold winter days, the Defender’s optional heated steering wheel provides comfort, allowing the driver to steer safely and comfortably before the vehicle heater warms up. The 4Runner doesn’t offer a heated steering wheel.
The Defender has a standard dual zone air conditioning allows the driver and front passenger to choose two completely different temperatures so people with different temperature preferences won’t have to compromise. This makes both the driver and front passenger as comfortable as possible. Dual zone air conditioning is only available on the 4Runner Limited/TRD Pro.
To quickly and conveniently keep personal devices charged without cables tangling and wearing out, the Land Rover Defender has a standard wireless phone charging system (Qi) in the center console. The 4Runner doesn’t offer wireless personal charging.
