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The Pathfinder offers a remote vehicle starting system, so the vehicle can be started from inside the driver's house. This allows the driver to comfortably warm up the engine before going out to the vehicle. The climate system will also automatically heat or cool the interior. The 4Runner doesn’t offer a remote starting system.
The engine computer on the Pathfinder automatically engages the starter until the car starts with one twist of the key and disables the starter while the engine is running. The 4Runner’s starter can be accidentally engaged while the engine is running, making a grinding noise and possibly damaging the starter and ring gear.
The Pathfinder Platinum’s standard easy entry system raises the steering wheel and glides the driver’s seat back when the door is unlocked or the ignition is switched off, making it easier for the driver to get in and out. The 4Runner doesn’t offer an easy entry system.
The Pathfinder Platinum has a standard heads-up display that projects speed, navigation instruction and incoming text message readouts in front of the driver’s line of sight, allowing drivers to view information without diverting their eyes from the road. The 4Runner doesn’t offer a heads-up display.
The Pathfinder’s power parking brake sets with one touch and releases with one touch or automatically. The 4Runner’s parking brake has to be released manually.
The power windows standard on both the Pathfinder and the 4Runner have locks to prevent small children from operating them. When the lock on the Pathfinder is engaged the driver can still operate all of the windows, for instance to close one opened by a child. The 4Runner prevents the driver from operating the other windows just as it does the other passengers.
The Pathfinder’s power window, power lock, power mirror and cruise control switches are lit from behind, making them plainly visible and easier to operate at night. The 4Runner’s power mirror and cruise control switches are unlit, making them difficult to find at night and operate safely.
The Pathfinder’s standard speed-sensitive wipers speed up when the vehicle does, so that the driver doesn’t have to continually adjust the speed of the wipers. The 4Runner’s manually variable intermittent wipers have to be constantly adjusted. The Pathfinder Platinum’s standard wipers adjust their speed and turn on and off automatically according to the amount of rainfall on the windshield.
To improve rear visibility by keeping the rear window clear, the Pathfinder has a standard rear fixed intermittent wiper with a full on position. The rear wiper standard on the 4Runner only has an intermittent setting, so in a hard rain visibility isn’t as good.
The Pathfinder has a standard automatic headlight on/off feature. When the ignition is on, the headlights automatically turn on at dusk and off after dawn. When the ignition turns off, the headlights turn off after a delay timed to allow you to securely get to your front door. The 4Runner has an automatic headlight on/off feature standard only on the Limited/TRD Pro.
Manual rear side window sunshades are available in the Pathfinder to help block heat and glare for the rear passengers. The 4Runner doesn’t offer rear side window sunshades.
The Pathfinder’s power mirror controls are mounted on the armrest for easy access. The 4Runner’s power mirror controls are on the dash where they are possibly hidden by the steering wheel and are awkward to manipulate.
When the Pathfinder Platinum is put in reverse, both rearview mirrors tilt from their original position. This gives the driver a better view of the curb during parallel parking maneuvers. Shifting out of reverse puts the mirrors into their original positions. The 4Runner’s mirrors don’t automatically adjust for backing.
Both the Pathfinder and the 4Runner offer available heated front seats. The Pathfinder also offers optional heated second row seats to keep those passengers extremely comfortable in the winter. Heated second row seats aren’t available in the 4Runner.
On extremely cold winter days, the Pathfinder SL/Platinum’s standard heated steering wheel provides comfort, allowing the driver to steer safely and comfortably before the vehicle heater warms up. The 4Runner doesn’t offer a heated steering wheel.
The Pathfinder has a standard dual zone air conditioning allows the driver and front passenger to choose two completely different temperatures so people with different temperature preferences won’t have to compromise. This makes both the driver and front passenger as comfortable as possible. Dual zone air conditioning is only available on the 4Runner Limited/TRD Pro.
To quickly and conveniently keep personal devices charged without cables tangling and wearing out, the Nissan Pathfinder (except S/SV) offers an optional wireless phone charging system (Qi) in the center console. The 4Runner doesn’t offer wireless personal charging.