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The Defender has a standard remote vehicle starting system, so the vehicle can be started from inside the driver's house. This allows the driver to comfortably warm up the engine before going out to the vehicle. The climate system will also automatically heat or cool the interior. The Land Cruiser doesn’t offer a remote starting system.
Unlike the driver-only memory system in the Land Cruiser, the Defender SE/HSE/X/V8 has standard driver and passenger memory, so that when drivers switch, the memory setting adjusts the driver’s seat, steering wheel position (with optional power wheel adjuster), outside mirror angle and climate settings and the front passenger seat also adjusts to the new passenger’s preset preferences.
The Defender (except Base) offers an available heads-up display that projects speed and navigation instruction readouts in front of the driver’s line of sight, allowing drivers to view information without diverting their eyes from the road. The Land Cruiser doesn’t offer a heads-up display.
The Defender’s power parking brake sets with one touch and releases with one touch or automatically. The Land Cruiser has a lever-type parking brake that has to be strenuously raised to engage properly. It has to be lifted up more and a button depressed to release it.
The power windows standard on both the Defender and the Land Cruiser have locks to prevent small children from operating them. When the lock on the Defender is engaged the driver can still operate all of the windows, for instance to close one opened by a child. The Land Cruiser prevents the driver from operating the other windows just as it does the other passengers.
Heated windshield washer nozzles are optional on the Defender to prevent washer fluid and nozzles from freezing and help continue to keep the windshield clear in sub-freezing temperatures. The Land Cruiser doesn’t offer heated windshield washer nozzles.
Consumer Reports rated the Defender’s headlight performance “Very Good,” a higher rating than the Land Cruiser’s headlights, which were rated “Good.”
The Defender’s power mirror controls are mounted on the armrest for easy access. The Land Cruiser’s power mirror controls are on the dash where they are possibly hidden by the steering wheel and are awkward to manipulate.
The Defender has standard heated front and optional heated second and third row seats, which keep the driver and passengers extremely comfortable in the winter. Toyota doesn’t offer heated seats in the third row of the Land Cruiser.
Optional air conditioned the front and second row seats keep the Defender’s passengers comfortable and take the sting out of hot leather in summer. The Land Cruiser doesn’t offer air-conditioned seats for the second row.